letters to nature Acknowledgements We thank M. Kobayashi, D. Engel, M. Muramatsu and T. Honjo for discussion and comments on the manuscript, M. Busslinger for information on RT–PCR analysis, N. Kaneko for the preparati

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• Transcription factors
• Molecular biology
• RNA
• SRG1 RNA
• GAL4/UAS system
• Upstream Activating Sequence
• General transcription factor
• Promoter
• TATA box
• Biology
• Gene expression
• Genetics